No, mature xylem cells do not have cytosol.
Mature xylem cells are specialized plant cells responsible for transporting water and minerals from the roots to the rest of the plant. During their development, these cells undergo a process called programmed cell death. This process results in the breakdown of the cell's organelles and the disappearance of the cytoplasm (which contains the cytosol). This leaves behind a hollow, reinforced cell wall, forming a continuous tube-like structure called a xylem vessel. This hollow structure allows for efficient water transport with minimal resistance. The lack of cytoplasm also prevents contamination of the water and nutrient stream.
Therefore, once xylem cells reach maturity and are fully functional in water transport, they are essentially dead cells lacking cytosol and other cellular components. Their primary role is structural support and fluid transport.
